Fixed #14588 -- Corrected r14393 to ensure that response middlewares are are always applied. Includes a battery of tests to validate the expected behavior of the middleware cycle. Thanks to Petr Marhoun for the report.

git-svn-id: http://code.djangoproject.com/svn/django/trunk@14398 bcc190cf-cafb-0310-a4f2-bffc1f526a37